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While historically thought of as simple leathery structures composed of skin, research has since shown that the wing membranes of pterosaurs were highly complex dynamic structures suited to an active style of flight.The outer wings (from the tip to the elbow) were strengthened by closely spaced fibers called actinofibrils.

Pterosaurs are often referred to in the popular media and by the general public as "flying dinosaurs", but this is scientifically incorrect.In some later pterosaurs, the backbone over the shoulders fused into a structure known as a notarium, which served to stiffen the torso during flight, and provide a stable support for the scapula (shoulder blade).Reconstructed wing planform of Quetzalcoatlus northropi (A) compared to the wandering albatross Diomedea exulans (B) and the Andean condor Vultur gryphus (C). northropi, the largest known flying animal, was more than three times as long as that of the wandering albatross.This membrane may have incorporated the first three fingers of the hand, as evidenced in some specimens.The brachiopatagium ("arm membrane") was the primary component of the wing, stretching from the highly elongated fourth finger of the hand to the hind limbs (though where exactly on the hind limbs it anchored is controversial and may have varied between species, see below).

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